
422 Many botanists have accepted this revision. In 2016, the Boraginales Working Group reevaluated the borage complex and recommended a separate waterleaf family. This is the system currently used by Jepson, 2 the authority for this Plant Guide. Subsequently, the availability of molecular data led to several reinterpretations of this relationship including the merging of the waterleaf family into the borage family. Species of Phacelia were previously placed in the waterleaf family (Hydrophyllaceae), based on morphological characteristics. Typical plants in the borage family are covered by small, stiff hairs, and have flowers that develop along a coiled stalk. The mature capsule is a dull brown and hairy, especially at top, and splits open to release up to four small seeds.īranching phacelia is a dicot angiosperm in the borage family (Boraginaceae). The seed capsule matures within the persistent calyx. Most flowers occur April through October. The style branches are slim, with minute stigmas the styles are white, turning lavender near the ends and remaining on the developing fruit. The pistil consists of one superior ovary, also covered with stiff hairs, There is a long forked style that is visible beyond the corolla and matures after the pollen has been shed. There are five long stamens clearly visible extending beyond corolla. Withered, brown petals remain on the flower stalk for some time, giving the entire structure an unkempt appearance. As the flower matures, the lobes reflex backward and the petals begin to turn brown. All-purple flowers are reported to occur elsewhere. The petals of our plants are mostly white or lavender-tinged, with five dark, purple spots in the throat. Flowers are bisexual and radially symmetrical with a deeply, five-lobed calyx and a five-lobed bell-shaped corolla. As the stalk unfurls, new flowers open at the apex of the flower while fruits develop along the straightened older stalk an older flower cluster often has the look of a fuzzy caterpillar. All parts of the plant are covered with stiff, erect, sharp hairs (trichomes), some of which are glandular, releasing a sticky exudate.įlowers are produced along a coiled stalk (a scorpioid cyme) with developing buds in the center of the coil. The larger leaves may be eight inches (20 cm) long, pinnately lobed or compound with the leaflets deeply, often irregularly, lobed or toothed. Although somewhat woody, the weak stems gain support by tangling with each other or from leaning on adjacent plants. The principle stems are up to five feet (0.5-1.5 m) long. … The wealth of information contained in this volume makes it an indispensable source for anybody in the fields of pure and applied plant sciences.Branching phacelia is a sprawling, untidy, perennial subshrub with several long branching stems from the base. "This volume, the seventh in this series, deals with 24 flowering plant families comprising a total of 911 genera. In conclusion, like all the previous volumes in the series, Volume VII of the Families and Genera of Vascular Plants provides highly useful, concise, and up-to-date treatments of what we presently know about the families dealt with." (Plant Systematics and Evolution) As in the previous volumes, all chapters meet high standards as to accuracy and accessibility of the information. Layout, printing, and binding of the book are esthetically applealing and of high quality. The editing was done by Joachim Kadereit, who obviously put in a great deal of effort to produce generally well balanced and consistently structured chapters. The contributions collected in the volume have been written by one or more internationally recognized specialists for the various families. "As with the earlier parts in the series, the long awaited Volume VII is an invaluable source of descriptive, taxonomic, phylogenetic, biogeographic, and last but not least the most critical bibliographic information.
